A study published in the Zoological Journal of the Linnean Society used molecular markers to identify all distinct lineages, their genetic variation and divergence times. As a result, it was revealed that phenotypical variation matches genetic diversity, with almost all subspecies corresponding to well-supported subclades. Three revealed lineages that diverged during the Plio-Pleistocene transition are proposed as different species, namely: Laudakia stellio (polytypic; Greece, Turkey, Lebanon), Laudakia cypriaca (monotypic; Cyprus) and Laudakia vulgaris (polytypic; Israel, Syria, Jordan, Egypt).
Reference
Karameta, E., Lymberakis, P., Grillitsch, H., Ilgaz, C., Avci, A., Kumlutaş, Y., Candan, K., Wagner, P., Sfenthourakis, S., Pafilis, P., Poulakakis, N. (2022). The story of a rock-star: multilocus phylogeny and species delimitation in the starred or roughtail rock agama, Laudakia stellio (Reptilia: Agamidae), Zoological Journal of the Linnean Society. Full-text here.